The presence of macaques in northern Africa dates back to the upper Miocene, about 6-7 million years ago. Shortly thereafter (about 5,5 million years ago), first representatives are found in Europe (Spain and Italy), which migrated on during the Messinian Salinity Crisis. Now, an article published in Journal of Human Evolution and led by research staff from the Institut Català de Paleontologia Miquel Crusafont (ICP), the Institut Català de Paleoecologia Humana i Evolució Social (IPHES-CERCA), the Universitat Rovira i Virgili

This is revealed by a new site, yielding the oldest stone tools documented to date in the Oldupai Gorge